1989 B.S., Kansas State University (KSU), KS
1991 D.V.M., Veterinary Medicine, KSU, KS
1991 M. Sc., Veterinary Pathology, KSU, KS
1995 D.Phil., Experimental Pathology, University of Oxford, Oxford, UK
1994-1995 E.P.A. Cephalosporin Junior Research Faculty
Lady Margaret Hall, University of Oxford, Oxford, UK
1995-2000 Assistant Professor, The Wistar Institute, Philadelphia, PA
1996-2000 Wistar Assistant Professor, Department of Pathobiology
University of Pennsylvania Veterinary School, PA.
2001-2006 Associate Professor, The Wistar Institute, Philadelphia, PA
2001-present Wistar Associate Professor of Microbiology
Department of Pathobiology, University of Pennsylvania Veterinary School, PA
2002-present Wistar Associate Professor of Medicine
Department of Medicine, University of Pennsylvania Medical School, PA
2007-present Professor, The Wistar Institute, Philadelphia, PA