Gavialis gangeticus, from the Indian subcontinent, is the only living gharial species related to the Puerto Rican gharial

28 million years ago "pepinianos", as the residents from he Puerto Rican town of San Sebastián are known, could frolic in the shores of their hometown. This is because 28 million years ago the now landlocked town of San Sebastián was a seashore town. Pepinianos looked very different then, though. There were no plazas, or traffic jams, or town fairs... as a matter of fact, 28 million years ago there were no humans, not in Puerto Rico, not anywhere else.

What San Sebasti·n did have were gharial: very large crocodiles basking in its shores.